Do any of the kiss lamps or the other column 2 lamps light when you single lamp test all the other lamps? This may prove whether there is a break in Column 2 which then is shorted to another column.

Interesting fault they work in attract mode and not in test mode. I'd be interested to know what others think. I would give the ASIC chip on the CPU a press to make sure it is seated properly or has a bad connection. Im not sure how the CPU controls lamps in attract mode compared to single lamp test.